With probably more detail than you'd like to know, this interesting New Yorker article talks about the origins of the #hashtag, pilcrow (¶), & ampersand, among others.
The Ancient Roots of Punctuation
The article is based on a new book by Keith Houston,
Shady Characters: The Secret Life of Punctuation, Symbols, and Other Typographical Marks. Sounds like a book every legal writing teacher should read!
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